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Thus, both signals 1 and 2 are required to achieve full T cell activation. Conversely, antigenic peptide stimulation (signal 1) in the absence of co stimulation (signal 2) cannot induce full T cell activation, thus resulting in T cell tolerance. In addition to co stimulatory molecules, there are also inhibitory molecules, such as CTLA 4 and PD 1, which induce negative signals to prevent T cell activation36,49.
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